Couple of questions on how this interacts with terrain and ignore cover rules?
If an opponent is shooting through dense does the +1 apply it would the unit need to be in terrain?
If a unit ignores light cover does the unit get +1 to save if it is in a terrain piece that grants light cover.
What about a piece of terrain that provides heavy cover?
1) It is considered to be receiving benefits of cover so will get the bonus but does not have to be within the terrain feature. Dense cover gives the bonus while the shots are taken through it, not into it.
2) Wording. NatCam works when unit is receiving benefits of cover. If an attack ignores those, then the NatCam bonus will not apply as the unit does not receive the benefits of cover for this attack.
3) It will work. Heavy Cover is still cover and unit within it will get the benefits od cover.
Sources:
Naturalised Camouflage
Each time a ranged attack is allocated to a model with this Adaptation while it is receiving the benefits of cover from a terrain feature, add an additional 1 to any armour saving throw made against that attack.
Terrain and Cover
Some pieces of terrain can provide certain bonuses to units in your army. Each bonus and how to acquire it is described in the Terrain Features section of the Core Rules. If a model has acquired such a bonus (to it's Save, Leadership or other characteristics), it is considered as receiving the benefit of cover. There are ways for some models to ignore such bonuses while making an attack, but it should be known what kind of benefit is ignored (see the Rare Rules for clarification).
BENEFITS OF COVER WHEN NOT IN TERRAIN
Sometimes a rule will tell you that a model or unit gains the benefit of cover, even while they are not entirely on or in a terrain feature. If a model or unit is under the effects of such a rule, and that rule does not specify what the benefits of cover are, when resolving an attack that model is assumed to be entirely on or within a terrain feature with the Light Cover terrain trait for all rules purposes. This means that when an attack made with a ranged weapon wounds a model under the effect of this rule, add 1 to the saving throw made against that attack (invulnerable saving throws are not affected).
If a model or unit has a rule that only applies while it is receiving the benefits of cover, then that rule will apply while that model or unit is under the effects of any rule that states it gains the benefit of cover even while they are not entirely on or within a terrain feature.
IGNORING THE BENEFITS OF COVER
Some rules allow a weapon or model to ignore the benefits that a target might otherwise receive from being on or in a terrain feature.
If a rule says that the target ‘does not receive the benefit of cover to its saving throw’, then, when resolving an attack with that rule, the models in the target unit ignore all benefits received from terrain traits that improve its saving throw (e.g. Light Cover, Heavy Cover).
If a rule says that the target ‘does not receive the benefits of cover that impose penalties on hit rolls’, then, when resolving an attack with that rule, models in the target unit ignore all benefits received from terrain traits that impose penalties on hit rolls (e.g. Dense Cover).
If a rule says that the target unit ‘does not receive the benefit of cover’, without specifying what benefits are ignored, then, when resolving an attack with that rule, models in the target unit ignore all benefits received from all terrain traits, including those that improve its saving throws, impose penalties on hit rolls and so on (e.g. Light Cover, Heavy Cover). Note that rules that say that the target unit ‘does not receive the benefit of cover’ do not enable a model or weapon with that rule to target a unit that would not normally be visible due to terrain features with the Obscuring trait.
IMPROVING THE BENEFITS OF COVER
Some models have abilities that improve the bonus a model receives to its saving throw when it is receiving the benefit of cover — for example, ‘add 2 instead of 1 to saving throws for models in this unit while it is receiving the benefit of cover’. If a model or unit with such a rule is receiving the benefits of cover from a terrain feature with the Light Cover or Heavy Cover trait, then this rule is applied as written. If that model or unit is receiving the benefits of cover from any other terrain features with any other terrain traits, you instead add 1 to its saving throws, in addition to any other benefits of cover that are gained from those terrain features. In either case, invulnerable saves are unaffected.